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By exploiting improvements in techniques for the isolation of intact spinach chloroplasts that retain the chloroplast envelope and therefore stromal components (as well as the thylakoid membranes), the authors were able to locate appreciable GR activity and contents of glutathione in this organelle

These factors contributed to the technique needing to be more sensitive than other assays

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Genetic complementation restored gGT expression (S2 Fig) and GSH-depleting activity (Fig 3A) to the gGT mutant