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INTRODUCTION Lifetime cigarette smoking has profound effects on health, affecting every organ in the body, and is causally associated with dozens of diseases and millions of deaths worldwide each year 1

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In order to investigate the direct effects of nicotine, Barr et al ., 2007 examined rat mesencephalic (midbrain) nerve cells using techniques such as enzyme mobility shift assays, immunoblot analysis, and kinase assays 26